Load ability definition before TryResolve so cooldown lookup cannot deny
after HP mutation (Bugbot). Happy Bruno cast uses beta; defeat spine keeps
alpha; bump spine seq and cooldown sleep margin for CI.
Update E5.M1 module doc, dependency register, and alignment table for
NEO-82 landed; Bruno defeat spine HP stepping; refactor four-pulse cast
test Arrange/Act/Assert; strike through resolved review items.
Invoke CombatOperations.TryResolve after E1 gates; nested combatResolution
on accept; per-ability catalog cooldown; target_defeated deny without
cooldown. Tests, Bruno defeat spine, README and module docs updated.